Here's an example of those holes. There are two layers of material on the upper, the mesh that you can see clearly, and a more fabric-like material underneath. The mesh got holes within a few months of purchasing them. They got bigger as I used them, and finally, the layer underneath the mesh tore as well. Right where the sole meets the upper.
Those shoes were so great while they lasted though.
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I got through 4 pairs of shoes whilst hiking the PCT this year, 2 of those pairs were Wildcats and I had exactly the same experience as you describe here. Really comfortable shoes but yeah, I got these holes appearing where the toebox flexes, usually after putting about 400 miles on them. In each case I continued to wear them for a couple hundred miles more but once the inner fabric began to fall apart, all kinds of dirt and dust was getting in and it was pretty annoying.
The other two pairs I wore through were Merrell's "All Out Charge" trail runners - I put 700 miles on one pair and 530 on another. Like the Wildcats they were very comfortable but the structure of the shoe on both pairs held up much better than the Wildcats - no holes at all. I found that the tread on the bottom, beneath the balls of my feet was the first thing to go. After about 500 miles it became, not really painful, but definitely noticeably less comfortable and this discomfort was the first indicator to me that I should think about swapping them out for a new pair. Overall I was very happy with the Merrell's.